@article {201, title = {Molecular phylogeny of the riffle beetle genus Hexanchorus revealed a presence of a new genus (Coleoptera: Elmidae)}, journal = { Arthropod Systematics \& Phylogeny}, volume = {80}, year = {2022}, pages = {575{\textendash}602}, abstract = {

The riffle beetle genus Hexanchorus Sharp is, with 25 known species, the most speciose genus of the subfamily Larainae in the Neotropics and the second largest globally. An analysis of its phylogeny, based on two mitochondrial (COI, 16S) and two nuclear (18S, 28S) markers, including Hexanchorus-like, but morphologically distinct specimens, supported presence of an unknown genus. The new genus, Rumilara gen. nov., is described here with four new species (R. obscura sp. nov., R. paterna sp. nov., R. riberai sp. nov., R. suppressa sp. nov.) and their larvae. The separate position of the new genus is, beside molecular differences, well substantiated by the morphology of adults (the absence of sexual dimorphism, structure of the pronotum) and larvae (pleurites never reaching to sixth abdominal ventrite). H. sagittatus stat. nov. is elevated to species rank based on molecular data, and Rumilara leleupi (Del\ève) comb. nov. is transferred from the genus Hexanchorus and redescribed, based on its morphology.

}, keywords = {Ecuador, Larainae, larva, molecular phylogeny, new genus, new species, Rumilara, South America, taxonomy}, doi = {10.3897/asp.80.e84013}, author = {M. Linsk{\'y} and Z. {\v C}iamporov{\'a}-Za{\v t}ovi{\v c}ov{\'a} and K. La{\v s}{\v s}ov{\'a} and F. {\v C}iampor Jr} } @article {197, title = {Okalia necopinata sp. nov. (Insecta, Coleoptera, Elmidae) from Gunung Mulu National Park in Sarawak (Malaysia)}, journal = {ZooKeys}, volume = {1092}, year = {2022}, pages = {79{\textendash}92}, abstract = {

Okalia necopinata sp. nov., from Sarawak, northwest Borneo, Malaysia, is described and illustrated along with an identifcation key. The standard barcoding fragment of the mitochondrial gene coding for cytochrome c oxidase subunit I (COI) was used together with morphological characters to delimit the taxonomic boundaries of the two known species, which live in shallow streams fowing through dense primary forests in limestone areas in Pahang (West Malaysia) and Sarawak (East Malaysia). The majority of all examined Okalia are fightless. Morphological distinguishing characters are the length of the granulated ffth elytral interval, the elytral and pronotal punctation, the aedeagal morphology, and the distal portion of the ovipositor.

}, keywords = {barcoding, Borneo, DNA, Dryopoidea, Macronychini, Oriental Region, rife beetles, taxonomy}, doi = {10.3897/zookeys.1092.79635}, author = {J. Kodada and M.A. J{\"a}ch and D. Selnekovi{\v c} and K. Goffov{\'a}} } @article {194, title = {A review of the Larainae of Australia with description of seven new species and the new genus Australara (Coleoptera, Byrrhoidea, Elmidae)}, journal = {ZooKeys}, volume = {1073}, year = {2021}, pages = {55{\textendash}117}, abstract = {

The three genera and four species of Larainae (Elmidae) previously described from Australia are reviewed, and one new genus and seven new species are described: Australara glaisteri gen. et sp. nov., Ovolara lawrencei sp. nov., Ovolara monteithi sp. nov., Stetholus carinatus sp. nov., Stetholus longipennis sp. nov., Stetholus metatibialis sp. nov., and Stetholus woronora sp. nov. A lectotype is designated for Hydora laticeps (Carter \& Zeck), and the first new collection records of the species are reported since its description in 1932. The occurrence in Australia of Potamophilinus papuanus Sat\ô, described from Papua New Guinea, is reported. A key to the species, photographic images of the external morphology and male genitalia, distribution maps, and habitat and behavioral information, when known, are provided for all twelve species of Australian Larainae.

}, keywords = {aquatic beetles, Hydora, new taxa, Ovolara, Potamophilinus, riffle beetles, Stetholus, taxonomy}, doi = {10.3897/zookeys.1073.71843}, author = {C.B. Barr and W.D. Shepard} } @article {193, title = {Zunielmis pax, a new genus and species of riffle beetles (Coleoptera: Elmidae) from Colombia}, journal = {Zoosystematica Rossica}, volume = {30}, year = {2021}, pages = {248{\textendash}265}, abstract = {

A new riffle beetle, Zunielmis pax gen. et sp. nov. , is described from Colombia (Vichada, Puerto Carre\ño, Bita River basin). It can be distinguished from all the other elmids by having large areas of a microgranular mesh (plastron) on the pronotum and elytra, crenulate lateral margin of the latter two, the shorter interocular distance than diameter of each eye, as well as by the characters of the structure of mandibles, the shape and sculpture of pronotum, and the structure ofmesosternum,mesotibiae and elytra. The imagoes were collected on freshwater sponges. The habitat, where the new species was found, is described, and the adaptive meaning of the characteristics of plastron are discussed. Drawings and photographs of the adult habitus, distinctive morphological characters, and male and female genitalia are provided.

}, keywords = {Coleoptera, Colombia, Elmidae, Elminae, freshwater sponges, Neotropics, new genus, new species, Orinoco River basin, riffle beetles, taxonomy}, doi = {10.31610/zsr/2021.30.2.248}, author = {M. Gonz{\'a}lez-C{\'o}rdoba and V. Manzo and C.E. Granados-Mart{\'\i}nez} } @article {192, title = {A new minute riffle beetle (Coleoptera: Elmidae: Pagelmis) from the Brazilian Amazon}, journal = {Zootaxa}, volume = {5061}, year = {2021}, pages = {369{\textendash}376}, abstract = {

The monotypic genus Pagelmis Spangler, 1981 was described from six thousand specimens of Pagelmis amazonica Spangler, 1981 collected with blacklight traps in Ecuador. For many years the known distribution of this genus was restricted to its type-locality, and recently it was recorded in Venezuela, Suriname and Brazil (Roraima State). Now, 40 years after the description of the genus, we describe the second Pagelmis species, based on male and female collected in the Amazon forest (Amazonas State, Brazil).With the description of the second species in the genus we provide an emended diagnosis of Pagelmis. Since adults of this genus were previously collected only in light traps, no habitat information was known. We collected adults of the new species in the first- and second-order streams with black, acidic water and sandy streambeds. The main substrates available in these streams were leaves and roots of riparian vegetation.

}, keywords = {aquatic insects, Elminae, emended diagnosis, Neotropical, taxonomy}, doi = {https://doi.org/10.11646/zootaxa.5061.2.9}, author = {T.T.S. Polizei and N. Hamada} } @article {191, title = {Amazonopsis cerdani (Coleoptera: Elmidae: Elminae), a New Species of Riffle Beetle from French Guiana with Habitat Observations}, journal = {The Coleopterists Bulletin}, volume = {75}, year = {2021}, pages = {427{\textendash}439}, abstract = {

A third species of Amazonopsis Barr, 2018, Amazonopsis cerdani Barr and Cerdan, new species (Coleoptera: Elmidae), is herein described from French Guiana. One female paratype of Amazonopsis theranyi Barr from Peru is tentatively reassigned to A. cerdani as a non-paratype. Photographic images of the male and female habitus, and the male genitalia, are provided, as is a distribution map and a key to the species.\ Amazonopsis cerdani differs from A. theranyi from Peru and Amazonopsis camachoi Barr from Venezuela by the presence of prominent spines on protarsomeres 1\–4 of males, among other characters. The habitat of this species is small, shallow, lowland streams with sandy-silty substrates and low flow. Specimens were collected from unconsolidated leaf litter in depositional areas, and from stick and leaf packs lodged in the current. Genetic analysis conducted on three specimens from two localities, a male and two females, showed that they are conspecific.

}, keywords = {aquatic beetles, DNA analysis, sexual dimorphism, South America, Streams, taxonomy}, doi = {10.1649/0010-065X-75.2.427}, author = {C.B. Barr and A. Cerdan and S. Clavier and J. Murienne} } @article {188, title = {Disersus otongachi a new species of Larainae riffle beetles from Ecuador (Coleoptera: Elmidae)}, journal = {Zootaxa}, volume = {4963}, year = {2021}, pages = {193{\textendash}199}, abstract = {

We describe here a new species in the genus Disersus Sharp, 1882 from the Otongachi Reserve in Ecuador. Disersus otongachi sp.nov. is externally similar to other representatives of the genus, however, this species can be clearly distinguished for significantly longer antennae and the unique shape of the male genitalia.

}, keywords = {DNA barcoding, Neotropical region, Otongachi Reserve, South America, taxonomy}, doi = {https://doi.org/10.11646/zootaxa.4963.1.12}, author = {F. {\v C}iampor Jr and J. Kodada and J. Boz{\'a}{\v n}ov{\'a} and Z. {\v C}iamporov{\'a}-Za{\v t}ovi{\v c}ov{\'a}} } @article {185, title = {A New, Sexually Dimorphic Species of Cylloepus Erichson from Brazil (Coleoptera: Elmidae)}, journal = {The Coleopterists Bulletin}, volume = {75}, year = {2021}, pages = {270{\textendash}274}, abstract = {

Cylloepus dimorphus Shepard, Sites, and Rodrigues, new species (Elmidae) is described from Brazil with a sexually dimorphic abdominal ventrite 5. Males have a protruding lobe on each side of the apex of ventrite 5. Females lack the lobes and are not distinguishable from congeners.

}, keywords = {Byrrhoidea, Neotropical, riffle beetle, taxonomy}, doi = {10.1649/0010-065X-75.1.270}, author = {W.D. Shepard and R.W. Sites and H.D.D. Rodrigues} } @article {183, title = {A revision of Onychelmis Hinton, 1941 (Coleoptera: Elmidae), with description of new species, DNA barcoding and notes on the geography of the genus}, journal = {European Journal of Taxonomy}, volume = {739}, year = {2021}, pages = {1{\textendash}35}, abstract = {

The genus Onychelmis Hinton, 1941 was for a long time regarded as a small taxon with only three known species distributed in the Andes. A study of new material from Ecuador, using morphological and molecular data, has resulted in the discovery of five new species: Onychelmis lenkae sp. nov., O. lobata sp. nov., O. minor sp. nov., O. onorei sp. nov. and O. splendida sp. nov. We also revised the entire genus and redescribed the three known species, O. longicollis (Sharp, 1882), O. leleupi Del\ève, 1968 and O. whiteheadi Spangler \& Santiago, 1991. Habitus photographs of adults are provided, together with line drawings of male and female genitalia, and schematic illustrations of the distribution of femoral tomentum for each species. DNA sequences for barcoding the COI mtDNA fragment were used to support species delimitation and to suggest possible relationships among species. The revision includes a key to adults of all species of Onychelmis and notes on the biogeography of the genus, with an updated distribution map.

}, keywords = {Andes, DNA barcoding, Elminae, South America, taxonomy}, doi = {https://doi.org/10.5852/ejt.2021.739.1263}, author = {M. Linsk{\'y} and Z. {\v C}iamporov{\'a}-Za{\v t}ovi{\v c}ov{\'a} and F. {\v C}iampor Jr} } @article {182, title = {Ancyronyx lianlabangorum sp. nov., a new spider riffle beetle from Sarawak, and new distribution records for A. pulcherrimus Kodada, J{\"a}ch \& {\v C}iampor based on DNA barcodes (Coleoptera, Elmidae)}, journal = {Zookeys}, volume = {1003}, year = {2020}, pages = {31{\textendash}55}, abstract = {

Ancyronyx lianlabangorum sp. nov. (Coleoptera, Elmidae), a new spider riffle beetle from the Kelabit Highlands (Sarawak, northern Borneo), is described. Illustrations of the habitus and diagnostic characters of the new species and the similar, polymorphic A. pulcherrimus Kodada et al. are presented. Differences to closely related species, based on COI nucleotide sequences and morphological characters, are discussed.\ Ancyronyx pulcherrimus is here recorded from Sarawak for the first time, based on DNA barcoding.

}, keywords = {Ancyronyx, Coleoptera, Elmidae, taxonomy, variability}, doi = {https://doi.org/10.3897/zookeys.1003.55541}, author = {J. Kodada and M.A. J{\"a}ch and H. Freitag and Z. {\v C}iamporov{\'a}-Za{\v t}ovi{\v c}ov{\'a} and K. Goffov{\'a} and D. Selnekovi{\v c} and F. {\v C}iampor Jr} } @article {181, title = {Notes on Stenhelmoides and description of the true male genitalia of S. strictifrons Grouvelle, 1908 (Coleoptera: Elmidae)}, journal = {Acta Amazonica}, volume = {50}, year = {2020}, pages = {317-326}, abstract = {

The genus Stenhelmoides comprises 15 species spread over all Central and South America and its greatest species richness is clearly associated with Amazonian regions. Based on examination of type specimens and additional material, we describe and illustrate the true male genitalia of Stenhelmoides strictifrons, which had until now been incorrectly illustrated in the literature. We also report new records of the genus Stenhelmoides from Brazil and Guyana. In addition to pictures of the species\’ habitus, we provide pictures of the type specimens of Stenhelmoides grandis, S. grouvellei, S. guyanensis; S. strictifrons and S. submaculus. Based on the International Code of Zoological Nomenclature, we recognize additional paralectotypes for S. guyanensis and for S. grouvellei.

}, keywords = {aquatic insects, morphology, Neotropics, riffle beetles, taxonomy}, doi = {https://doi.org/10.1590/1809-4392202002791}, author = {A.S. Fernandes and T.T.S. Polizei and R. Boldrini} } @article {165, title = {Hexacylloepus Hinton, 1940 rises again; 18 new species of a neglected genus (Coleoptera: Byrrhoidea: Elmidae)}, journal = {Zootaxa}, volume = {4819}, year = {2020}, pages = {006{\textendash}048}, abstract = {

The genus Hexacylloepus extends from the Neotropical region to the southwestern United States of America. At present, 25 species are known from 20 countries, with no new species proposed since 1972. In this paper, we describe and illustrate 18 new species (H. barrae; H. calori; H. casariae; H. froehlichi; H. geiseri; H. grandis; H. iassu; H. keitai; H. maierae; H. manauara; H. metapa; H. phalluspilosus; H. shorti; H. taylorae; H. thoracica; H. tibialis; H. ubirajarai; H. zaninii), from eight countries and give the first records of the genus from Suriname and Guyana.

}, keywords = {aquatic beetles, morphology, Neotropical, riffle beetles, taxonomy}, doi = {https://doi.org/10.11646/zootaxa.4819.1.2}, author = {T.T.S. Polizei and M.V.L. Barclay and P.d.C. Bispo} } @article {164, title = {The Neotropical genus Portelmis Sanderson, 1953 (Coleoptera: Elmidae): three new species, new records and updated key}, journal = {Zootaxa}, volume = {4810}, year = {2020}, pages = {452{\textendash}467}, abstract = {

Elmidae is one of the most diverse families of aquatic beetles, with more than 1500 species in 151 genera. The Neotropical genus Portelmis formerly comprised five species known from Central and northern South America, divided into two species groups. Three new Brazilian species from the nevermanni group are described and illustrated here: Portelmis vanini sp. nov., Portelmis krolowi sp. nov. and Portelmis fusariae sp. nov. New species records for the North, Central-West and Southeast of Brazil are presented.We also provide first SEM images and an updated key for Portelmis species.

}, keywords = {aquatic insects, riffle beetles, South America, taxonomy}, doi = {https://doi.org/10.11646/zootaxa.4810.3.3}, author = {T.T.S. Polizei and A.S. Fernandes} } @article {163, title = {Unexpected discovery of a new Podonychus species in Kyushu, Japan (Coleoptera, Elmidae, Elminae, Macronychini)}, journal = {Zookeys}, volume = {933}, year = {2020}, pages = {107{\textendash}123}, abstract = {

Podonychus gyobu sp. nov., a second species of the genus Podonychus J\äch \& Kodada, 1997, hitherto known only from Indonesia, is described from Kyushu, Japan. This new species is similar to P. sagittarius\ J\äch \& Kodada, 1997, but differs from it in the straight penis, arcuate 2nd labial palpomere, and in the 3rd antennomere being longer than wide. The endophallic structures and the larva of P. gyobu sp. nov. are described. A character matrix of the Macronychini genera and a key to the Japanese genera are provided.

}, keywords = {disjunct distribution, endophallus, larvae, new species, riffle beetle, SEM, taxonomy}, doi = {https://doi.org/10.3897/zookeys.933.48771}, url = {https://zookeys.pensoft.net/article/48771/}, author = {H. Yoshitomi and M. Hayashi} } @article {162, title = {Austrolimnius cleidecostae, a new species of riffle beetle (Coleoptera: Elmidae) from Brazil with habitat notes and updated key of the Brazilian species of the genus}, journal = {Pap{\'e}is Avulsos de Zoologia}, volume = {60}, year = {2020}, pages = {1-10}, abstract = {

Austrolimnius cleidecostae sp. nov. is one of the smallest species of Elmidae from the Neotropical region. The new species is described and illustrated based on male and female specimens. This species represents the first record in Elmidae of mandibles with an articulated, sclerotized process on inner subapical edge. The specimens were collected from streams in the Atlantic rainforest of S\ão Paulo and Santa Catarina, and habitat notes are given. An updated key for the Austrolimnius Carter \& Zeck, 1929 of Brazil is provided. Austrolimnis browni Hinton, 1971 and A. musgravei Hinton, 1939 are recorded for the first time from Brazil.

}, keywords = {aquatic beetles, Neotropical, South America, Streams, taxonomy}, doi = {http://doi.org/10.11606/1807-0205/2020.60.special-issue.12}, author = {T.T.S. Polizei and L.d.S.M. Costa and P.d.C. Bispo} } @article {160, title = {Additional new species of Grouvellinus Champion 1923 (Insecta, Coleoptera, Elmidae) discovered by citizen scientists and DNA barcoded in the field applying a novel MinION-based workflow}, journal = {Journal of Natural History}, volume = {53}, year = {2020}, pages = {2593-2620}, abstract = {

Four new species, Grouvellinus nigerquadratus sp. nov., G. cruxniger sp. nov., G. luciaensis sp. nov., and G. borneensis sp. nov., are described from Borneo and illustrated in detail. Two additional species of the genus which remain undescribed are briefly diagnosed. The material was collected during Taxon Expeditions\’ field course which involved citizen scientists, students and taxonomists. Specimens were collected at altitudes between 270 m and 750 m above sea level using fine-meshed hand-nets and blacklight traps. The morphological species delimitation based on morphology was supplemented and congruent with mtDNA sequences which are the first DNA barcodes for the genus from Borneo. They were obtained in the field using a newly developed rapid and accurate MinION-based workflow. The inter \– and intraspecific genetic distances and the problems regarding cryptic species delimitation are discussed.

}, keywords = {Citizen science, nanopore sequencing, riffle beetle, taxon expeditions, taxonomy}, doi = {https://doi.org/10.1080/00222933.2019.1709669}, author = {H. Freitag and C. Molls and A.M. Bouma and J.M. Garces and M. Rossato and E. Cosentino and M. Delledonne} } @article {159, title = {A new species of the genus Graphelmis DEL{\`E}VE, 1968 from China (Coleoptera: Elmidae)}, journal = {Zootaxa}, volume = {4728}, year = {2020}, pages = {483{\textendash}488}, abstract = {

Graphelmis dulongensis sp. nov. is described from Yunnan, China. Photos of habitus and male genitalia are given.

}, keywords = {China, new species, riffle beetles, taxonomy, Yunnan}, doi = {https://doi.org/10.11646/zootaxa.4728.4.7}, author = {X. Dong and D. Bian} } @article {158, title = {A new species of Macrelmis Motschulsky, 1859 (Coleoptera: Elmidae) and new records of Elmidae from Roraima State, northern Brazil}, journal = {Zootaxa}, volume = {4718}, year = {2020}, pages = {277{\textendash}284}, abstract = {

A new species of Macrelmis Motschulsky, 1859 (Macrelmis rodrigoi sp. nov.) is described and illustrated based on specimens found in mountainous areas of the northern Roraima State, Brazil. Pagelmis amazonica Spangler, 1981, Potamophilops bostrychophallus Maier, 2013 and Stegoelmis geayi Spangler, 1990 are recorded in Brazil for the first time and new records for Gyrelmis obesa Hinton, 1940, Macrelmis thorpei Hinton, 1945 and Neolimnius palpalis Hinton, 1939 are presented.

}, keywords = {aquatic insects, entomology, rifle beetles, South America, taxonomy}, doi = {https://doi.org/10.11646/zootaxa.4718.2.9}, author = {M.d.L.S.d. Almeida and A.S. Fernandes and R. Boldrini} } @article {154, title = {The genus Hintonelmis (Coleoptera: Elmidae: Elminae), new species and records}, journal = {Journal of Natural History}, volume = {52}, year = {2019}, pages = {2949-2959}, abstract = {

Hintonelmis is a South American genus of Elmidae, comprising 11\ species. Two further new species are here described and illustrated: Hintonelmis guianensis sp. nov. from Guyana, and Hintonelmis roellae sp. nov. from Brazil, can be distinguished from all other known members of the genus by their general colour pattern, some characters of pronotal and elytral sculpture, and the morphology of the male genitalia. The type material is deposited in the Natural History Museum,\ London, UK and Museum of Zoology of the University of S\ão Paulo, S\ão Paulo, Brazil. New distributional data for species of Hintonelmis are\ presented for the South of Brazil, Bolivia, Ecuador, and Guyana. We also\ provide an updated key for the genus species.

}, keywords = {aquatic beetles, Neotropical, riffle beetles, South America, taxonomy}, doi = {https://doi.org/10.1080/00222933.2018.1560512}, author = {T.T.S. Polizei and M.V.L. Barclay} } @article {153, title = {The genus Cylloepus Erichson, 1847 (Coleoptera: Elmidae: Elminae) new species and combinations}, journal = {Zootaxa}, volume = {4652}, year = {2019}, pages = {93{\textendash}100}, abstract = {

Elmidae is one of the most diverse families of aquatic beetles, with more than 1500 species in 148 genera. Cylloepus is the genus with the most species in Americas, with 52 species and 2 subspecies currently known. The genus is easily recognised by external morphology; body elongate, pronotum with a longitudinal impression on the disc and elytra striate and punctate. In this paper, based on the study of several international reference collections, we propose two new combinations, and describe two new species of Cylloepus (C. bispoi sp. nov. and C. segurae sp. nov.) from Ecuador and Bolivia.

}, keywords = {aquatic beetles, Neotropical region, riffle beetles, South America, taxonomy}, doi = {https://doi.org/10.11646/zootaxa.4652.1.3}, author = {T.T.S. Polizei and M.V.L. Barclay} } @article {152, title = {Macrelmis pubescens (Grouvelle, 1889) {\textendash} redescription of the species, designation of a lectotype and description of its larva (Coleoptera: Elmidae)}, journal = {Annales de la Soci{\'e}t{\'e} entomologique de France (N.S.)}, year = {2019}, abstract = {

The adult of Macrelmis pubescens (Grouvelle, 1889) is redescribed, illustrated, a lectotype designated, and new locality information provided. The lectotype is housed in the Grouvelle\’s collection in the Mus\éum national d\’Histoire naturelle, Paris, France. The larva of M. pubescens is described, illustrated and compared with the known larvae of the genus. A diagnosis for Macrelmis Motschulsky, 1860 larvae is provided.

}, keywords = {aquatic beetles, Brazil, Neotropical, riffle beetles, South America, taxonomy}, doi = {https://doi.org/10.1080/00379271.2019.1610669}, author = {T.T.S. Polizei and S.A. Casari} } @article {149, title = {Amazonopsis, an unusual new genus of riffle beetle from South America with two new species (Coleoptera, Elmidae, Elminae)}, journal = {ZooKeys}, volume = {803}, year = {2018}, pages = {71{\textendash}92}, abstract = {

Amazonopsis gen. n. is described to include A. theranyi sp. n. from Peru, Venezuela and French Guiana, and A. camachoi sp. n. from Venezuela. The descriptions are accompanied by figures illustrating the male and female habitus of A. theranyi, the male habitus of A. camachoi, and male genitalia of both species. Amazonopsis theranyi exhibits pronounced secondary sexual dimorphism which likewise may be a characteristic of the genus. Amazonopsis males have modifed protarsal and mesotarsal claws, a pair of small spines on the anterior prosternum, and a pair of ventrally directed processes on the posterior metaventrite. Females of A. theranyi display a pair of unique, oval perforations in the cuticle of the pronotum and have unmodifed claws; females of A. camachoi are unknown. Descriptions are furnished of the stream habitats and microhabitats where the study specimens were collected.

}, keywords = {aquatic beetles, French Guiana, Neotropics, peri-Amazonian, Peru, sexual dimorphism, taxonomy, Venezuela}, doi = {10.3897/zookeys.803.28124}, author = {C.B. Barr} } @article {140, title = {Dryopomorphus siamensis sp. nov., a new riffle beetle from Thailand (Coleoptera: Elmidae) and remarks on the morphology of the mouthparts and hind wing venation of D. bishopi Hinton}, journal = {Entomological Problems}, volume = {24}, year = {1993}, pages = {51{\textendash}58}, abstract = {

Dryopomorphus siamensis\ sp. nov.\ from Thailand is described. Line drawings of the antenna and the male genitalia are provided. In addition, the labrum, mandible, maxilla, labium and the hind wing of\ D. bishopi\ are examined and illustrated.

}, keywords = {Coleoptera, Dryopomorphus, Elmidae, hind wing venation, morphology, mouthparts, new species, taxonomy}, author = {J. Kodada} } @article {138, title = {Two new species of the genus Dryopomorphus Hinton, 1936 from China (Coleoptera, Elmidae)}, journal = {ZooKeys}, volume = {765}, year = {2018}, pages = {51{\textendash}58}, abstract = {

Two new species of the genus Dryopomorphus Hinton, 1936 are described from China: Dryopomorphus heineri sp. n. and D. ruiliensis sp. n. Habitus photographs and detailed line drawings of the male genitalia are provided.

}, keywords = {Dryopomorphus, Guangdong, riffle beetle, taxonomy, Yunnan}, doi = {10.3897/zookeys.765.24366}, url = {https://zookeys.pensoft.net/articles.php?id=24366}, author = {D. Bian and X. Dong and Y. Peng} } @article {137, title = {Dryopomorphus hendrichi sp.nov. from West Malaysia (Coleoptera: Elmidae)}, journal = {Entomological Problems}, volume = {36}, year = {2006}, pages = {71-73}, abstract = {

Dryopomorphus hendrichi sp.nov. from Johor, West Malaysia is described, illustrated and compared with known Dryopomorphus species.

}, keywords = {Coleoptera, Dryopomorphus, Elmidae, new species, Southeast Asia, taxonomy}, author = {F. {\v C}iampor Jr and J. Kodada} } @article {132, title = {Microcylloepus loebli, a new species of riffle beetle (Coleoptera, Elmidae)}, journal = {Entomologische Bl{\"a}tter und Coleoptera}, volume = {114}, year = {2018}, pages = {323-327}, abstract = {

Microcylloepus loebli n. sp. represents the first record of a riffle beetle from Bahia State, Brazil. The new species is described and illustrated, and the type material is deposited in the Museum of Zoology of the University of S\ão Paulo, S\ão Paulo, Brazil and the Natural History Museum, London, UK.

}, keywords = {aquatic beetles, Neotropical fauna, taxonomy}, author = {T.T.S. Polizei} } @article {83, title = {Review of the Korean Elmidae (Coleoptera: Dryopoidea) with descriptions of three new species}, journal = {Aquatic Insects}, volume = {36}, year = {2014}, pages = {93-124}, abstract = {

The Korean species of the riffle beetle family Elmidae are revised. Thirteen species belonging to nine genera are recognized including three new species, Grouvellinus aerosus sp. n., Zaitzeviaria kyungseoki sp. n., and Z. obesa sp. n., and one unidentified species of Ordobrevia Sanderson, 1953. Leptelmis coreana Jung and Bae, 2012 and\ L. ochra Jung and Bae, 2012 are synonymized with L. gracilis gracilis Sharp, 1888. Four species, Macronychus levanidovae Lafer, 1980, Stenelmis koreana Sat\ô, 1978, S. cf. montana Zhang and Yang, 1995, and Zaitzevia tsushimana Nomura, 1963, are newly recorded from South Korea. Descriptions, type information, synonymies, illustrations of diagnostic characters and geographical variation, a checklist, and a key to the adults of the Korean species are provided.

}, keywords = {Coleoptera, Elmidae, Korea, new faunistic records, new species, new synonym, taxonomy}, author = {S.W. Jung and M.A. J{\"a}ch and Y.J. Bae} } @article {79, title = {A new species of Stenelmis (Coleoptera: Elmidae) from South India}, journal = {International Journal of Environmental Biology}, volume = {6}, year = {2016}, pages = {24-28}, abstract = {

A new species of riffle beetle, Stenelmis pandiani sp.n. is described on the basis of male and female adults collected from South India. The morphological and molecular data confirm species assignment and distinguish as a new species.

}, keywords = {DNA barcoding, new species, riffle beetle, taxonomy}, author = {S. Anbalagan and M. Kannan and K. Rekha and M. Krishnan and G. Ponraman and S. Dinakaran} } @article {69, title = {Description and notes on the bionomics of a new species of Potamophilops Grouvelle, 1896 (Coleoptera: Elmidae: Larainae), from the Cerrado biome in Brazil}, journal = {Journal of Natural History}, volume = {46}, year = {2012}, pages = {717{\textendash}727}, abstract = {

Potamophilops bragaorum sp. nov. is described and illustrated based on adult specimens collected in a mountainous area in the Cerrado biome in Taquaru\ç\ú district, Tocantins state, Brazil. This is the second species described in the genus and it represents the northernmost record of Potamophilops, which is known only from Argentina and Brazil. In this article we provide a diagnosis, details on the morphology and images of the habitus and structures used to determine the species and to distinguish the gender. Field observations permitted the description of some aspects on the biology of the new species that seem to be very similar to those of other Larainae genera. Examination of gut contents revealed a diet based mostly on periphyton.

}, keywords = {aquatic insects, Cerrado biome, Neotropics, riffle beetles, taxonomy}, author = {A.S. Fernandes and N. Hamada} } @article {67, title = {The genus Portelmis Sanderson, 1953 (Coleoptera: Elmidae: Elminae): first report in Brazil, description of two new Amazonian species and species key for males}, journal = {Zootaxa}, volume = {2517}, year = {2010}, pages = {33-43}, abstract = {

The genus Portelmis Sanderson, 1953 and the species P. gurneyi Spangler, 1980 are reported for the first time in Brazil. Portelmis kinonatilis sp. nov. and P. paulicruzi sp. nov. are described and illustrated based on adult specimens collected in two different municipalities in the state of Amazonas, Brazil. Diagnoses and illustrations of all Portelmis species, and a taxonomic key to identify males are presented.

}, keywords = {aquatic insects, kinon, Neotropics, Portelmis, riffle beetle, taxonomy}, author = {A.S. Fernandes and M.I.S. Passos and N. Hamada} } @article {66, title = {Two new species of Austrolimnius Carter \& Zeck (Insecta: Coleoptera: Elmidae) from Southeastern Brazil}, journal = {Zootaxa}, volume = {3389}, year = {2012}, pages = {14-24}, abstract = {

The genus Austrolimnius Carter \& Zeck has its greatest known diversity in Oceania, but its fauna in the Americas is poorly known. In this paper, two new species\—Austrolimnius uncatus sp. nov. and A. bocainensis sp. nov.\—are described and illustrated based on adult specimens collected at Serra da Bocaina National Park, S\ão Jos\é do Barreiro Municipality, S\ão Paulo State, and Nova Friburgo Municipality, Rio de Janeiro State, Brazil. Austrolimnius uncatus sp. nov. is characterized by the long setae on the mesotarsus, the medial expansion of the hind tibiae, and the hook-like shape of the male genitalia. Austrolimnius bocainensis sp. nov. can be recognized by the absence of setae on the mid-tarsus, the cylindrical hind tibiae, and the slightly curved genitalia with few setae. A pictorial key for the Brazilian species of the genus is given. New records of Brazilian species of the genus are reported.

}, keywords = {aquatic insects, Atlantic forest, Elminae, taxonomy}, author = {G.S. Miranda and B.H.L. Sampaio and M.I.S. Passos} } @article {63, title = {Two new species of Cuspidevia J{\"a}ch \& Boukal, 1995 from China (Coleoptera: Elmidae)}, journal = {Zootaxa}, volume = {2663}, year = {2010}, pages = {53-58}, abstract = {

Two new species of the genus Cuspidevia J\äch \& Boukal, 1995 (Macronychini) are reported from Jiangxi Province,
China: Cuspidevia brevis sp. nov., Cuspidevia jaechi sp. nov. Descriptions, diagnoses, illustrations and a key to
Cuspidevia are given.

}, keywords = {China, Cuspidevia, Macronychini, new species, taxonomy}, author = {D. Bian and L. Ji} } @article {59, title = {Typhloelmis Barr (Coleoptera: Elmidae: Elminae), a New Stygobiontic Riffle Beetle Genus with Three New Species from Texas, USA}, journal = {The Coleopterists Bulletin}, volume = {69}, year = {2015}, pages = {531-558}, abstract = {

The adult and larva of Typhloelmis Barr, a new genus of eyeless, stygobiontic elmid from springs in west-central
Texas, and the adults of three new species, Typhloelmis caroline Barr, Typhloelmis finegan Barr, and Typhloelmis sanfelipe Barr, are described, diagnosed, and illustrated. Taxonomic couplets are provided for the insertion of the new genus intoexisting adult elmid generic keys, and an adult key to the species of Typhloelmis is included. Stygobiontic elmids are reviewed, and the uniqueness of Typhloelmis is discussed. Typhloelmis is the first stygobiontic elmid genus reported from the Nearctic ecozone and is the seventh described worldwide. It is only the second elmid genus known to be anophthalmic (completely eyeless). The Edwards-Trinity Aquifer and the spring habitat in which the genus and species occur are described in detail, and potential threats are discussed.

}, keywords = {aquatic beetles, Edwards-Trinity Aquifer, karst spring, larva, stygofauna, taxonomy}, doi = {http://dx.doi.org/10.1649/0010-065X-69.4.531}, url = {http://www.bioone.org/doi/full/10.1649/0010-065X-69.4.531}, author = {C.B. Barr}, editor = {J.R. Gibson and P.H. Diaz} } @article {49, title = {A taxonomic review of the Oriental species of the genus Ancyronyx Erichson, 1847 (Coleoptera, Elmidae)}, journal = {Revue suisse de Zoologie}, volume = {101}, year = {1994}, pages = {601-622}, chapter = {601}, abstract = {

The Oriental species of the genus\ Ancyronyx\ Erichson are revised. It is shown that\ A. acaroides\ Grouvelle is correctly placed in the genus\ Ancyronyx.\ Five new species and one new subspecies are described:\ Ancyronyx acaroides cursor\ ssp. n. (Bali),\ A. johanni\ sp. n. (Siberut),\ A. malickyi\ sp. n. (Sumatra, Thailand, Borneo),\ A. procerus\ sp. n. (Borneo),\ A. sarawacensis\ sp. n. (Borneo) and\ A. schillhammeri\ sp. n. (Mindoro). A lectotype is designated for\ A. acaroides.\ A key to the world species is included.\ Ancyronyx constrictus\ Reitter is transferred to the genus\ Stenelmis\ Dufour.

}, keywords = {Ancyronyx, Coleoptera, Elmidae, Oriental Region, taxonomy}, author = {M.A. J{\"a}ch} } @article {46, title = {Larvae and a new species of Ancyronyx Erichson, 1847 (Insecta, Coleoptera, Elmidae) from Palawan, Philippines, using DNA sequences for the assignment of the developmental stages}, journal = {Zookeys}, volume = {136}, year = {2011}, pages = {47-82}, abstract = {

Ancyronyx montanus sp. n. is described based on adults and larvae, matched using their cox1 DNA sequence data. Larvae of six additional species of Ancyronyx Erichson, 1847 were also described here for the first time, aided by cox1 or cob data: A. helgeschneideri A. helgeschneideri Freitag \& J\äch, 2007, A. minerva Freitag \& J\äch, 2007, A. patrolus Freitag \& J\äch, 2007, A. procerus J\äch, 1994, A. punkti Freitag \& J\äch, 2007, A. pseudopatrolus Freitag \& J\äch, 2007. Ancyronyx procerus is newly recorded from the Philippines by a larval specimen from Busuanga island. The new species and larval stages are described in detail and illustrated by digital and SEM images. A key to the Ancyronyx larvae of Palawan and an updated checklist of Philippine Ancyronyx is provided.

}, keywords = {Ancyronyx, AQUA Palawana, cox1, DNA barcoding, Elmidae, larva, new species, Spider Water Beetle, taxonomy}, doi = {10.3897/zookeys.136.1914}, author = {H. Freitag and M. Balke} } @article {45, title = {The genus Ancyronyx Erichson, 1847 (Coleoptera, Elmidae) in Palawan and Busuanga, (Philippines) with descriptions of six new species}, journal = {Zootaxa}, volume = {1590}, year = {2007}, pages = {37-59}, abstract = {

The species of the elmid genus Ancyronyx Erichson from the Philippine islands of Palawan and Busuanga are revised. Six new species are described: Ancyronyx helgeschneideri Freitag \& J\äch sp. n., A. minerva Freitag \& J\äch sp. n., A. minutulus Freitag \& J\äch sp. n., A. patrolus Freitag \& J\äch sp. n., A. pseudopatrolus Freitag \& J\äch sp. n., and A. punkti Freitag \& J\äch sp. n. A checklist and a key to the species of Ancyronyx recorded from the Philippines are provided.

}, keywords = {Ancyronyx, Busuanga, Coleoptera, Elmidae, Palawan, Philippines, Southeast Asia, taxonomy, water beetles}, author = {H. Freitag and M.A. J{\"a}ch} } @article {44, title = {Ancyronyx Erichson, 1847 (Coleoptera, Elmidae) from Mindoro, Philippines, with description of the larvae and two new species using DNA sequences for the assignment of the developmental stages}, journal = {Zookeys}, volume = {321}, year = {2013}, pages = {35-64}, abstract = {

Ancyronyx buhid sp. n. and A. tamaraw sp. n. are described based on adults and larvae, matched using their cox1 or cob DNA sequence data. Additional records of A. schillhammeri J\äch, 1994 and A. minerva Freitag \& J\äch, 2007 from Mindoro are listed. The previously unknown larva of A. schillhammeri is also described here, aided by cox1 data. The new species and larval stages are described in detail and illustrated by SEM and stacked microscopic images. Keys to the adult and larval Ancyronyx species of Mindoro and an updated checklist of Philippine Ancyronyx species are provided. The usefulness as bioindicators, the phylogenetic relationships and biogeographic aspects affecting the distribution patterns are briefy discussed.

}, keywords = {Ancyronyx, Buhid, cob, Coleoptera, cox1, DNA barcoding, Elmidae, larva, Mindoro, new species, Philippines, Spider Water Beetles, taxonomy}, doi = {10.3897/zookeys.321.5395}, author = {H. Freitag} } @article {32, title = {A new species of Hintonelmis Spangler (Coleoptera: Elmidae: Elminae) from Central Amazonia, Brazil}, journal = {Zootaxa}, volume = {2353}, year = {2010}, pages = {43-48}, chapter = {43}, abstract = {

Hintonelmis anamariae sp. nov. is described and illustrated based on adult specimens collected in Presidente Figueiredo Municipality, Amazonas State, Brazil. This species can be distinguished from all other known Hintonelmis species by its dorsal color pattern and the morphology of the male genitalia.

}, keywords = {Elmidae, morphology, Neotropics, riffle beetle, taxonomy}, author = {A.S. Fernandes and M.I.S. Passos and N. Hamada} } @article {29, title = {Malaysian species of Dryopomorphus Hinton, 1936 (Insecta: Coleoptera: Elmidae)}, journal = {Zootaxa}, volume = {3564}, year = {2012}, pages = {1-16}, abstract = {

We provide a review of species of the genus Dryopomorphus Hinton, 1936 occurring in Malaysia. Three known species: D. bishopi Hinton, 1971; D. satoi Spangler, 1985 and D. hendrichi\ {\v C}iampor Jr \& Kodada, 2006 are diagnosed and five new species from the Malay Peninsula and Borneo are described here: D. grandis, D. jaechi, D. memei, D. pekariki and D. sarawacensis. Habitus photographs, drawings of genitalia and diagnostic characters of all species are included.

}, keywords = {Coleoptera, Dryopomorphus, Elmidae, new species, Southeast Asia, taxonomy}, author = {F. {\v C}iampor Jr and Z. {\v C}iamporov{\'a}-Za{\v t}ovi{\v c}ov{\'a} and J. Kodada} } @article {22, title = {Systematic revision of the genus Graphelmis (Coleoptera: Elmidae) VIII. Three new species from the Graphelmis marshalli species group and distributional notes on G. grouvellei}, journal = {Entomological Problems}, volume = {36}, year = {2006}, pages = {13-20}, chapter = {13}, abstract = {

{\v C}IAMPOR Jr., F. 2005. Systematic revision of the genus Graphelmis (Coleoptera: Elmidae) VIII. Three new species from the Graphelmis marshalli species group and distributional notes on G. grouvellei. Entomol. Probl. 36(1): 13\–20. \– Additional three new species of the genus Graphelmis DEL\ÈVE, 1968 are described from Indonesia and Malaysia: G. anulata sp.nov., G. merkli sp.nov. and G. hintoni sp.nov., all belonging to the G. marshalli species group. Habitus and detailed drawings of characteristic structures are given and differential diagnosis of the species within the G. marshalli species group is provided. The additional material of G. grouvellei DEL\ÈVE, 1970 was revised, and the species is recorded from Malaysia for the first time.

}, keywords = {Coleoptera, distribution, Elmidae, Graphelmis, Southeast Asia, taxonomy}, author = {F. {\v C}iampor Jr} } @article {21, title = {Systematic revision of the genus Graphelmis (Coleoptera: Elmidae) VII. Graphelmis obesa species group}, journal = {Entomological Problems}, volume = {35}, year = {2005}, pages = {117-122}, chapter = {117}, abstract = {

{\v C}IAMPOR Jr., F. 2005. Systematic revision of the genus Graphelmis (Coleoptera: Elmidae) VII. Graphelmis obesa species group. Entomol. Probl. 35(2): 117\–122. \– The ninth new species group is designated within the genus Graphelmis DEL\ÈVE, 1968, including two new species: G. obesa sp.nov. from Sabah and G. gemuk sp.nov. from Sarawak (both Borneo, Malaysia). Habitus and detailed drawings of characteristic structures of genitalia and pregenital segments of males are given.

}, keywords = {Coleoptera, Elmidae, Graphelmis, Southeast Asia, taxonomy}, author = {F. {\v C}iampor Jr} } @article {20, title = {Systematic revision of the genus Graphelmis (Coleoptera: Elmidae) VI. Graphelmis marshalli species group}, journal = {Entomological Problems}, volume = {35}, year = {2005}, pages = {11-38}, chapter = {11}, abstract = {

{\v C}IAMPOR Jr., F. 2005. Systematic revision of the genus Graphelmis (Coleoptera: Elmidae) VI. Graphelmis marshalli species group. Entomol. Probl. 35(1): 11\–38. \– The sixth new species group Graphelmis marshalli is designated within the genus Graphelmis DELEVE, 1968, comprising six known and ten new species: G. bouchardi (GROUVELLE, 1896), G. consobrina DELEVE, 1968, G. dentipes sp.nov., G. elegans sp.nov., G. fuscicornis sp.nov., G. grouvellei DELEVE, 1970, G. schoedli sp.nov., G. malickyi sp.nov., G. marshalli (HINTON, 1936), G. monticola (GROUVELLE, 1896), G. nitida sp.nov., G. quadrimaculata sp.nov., G. robusta sp.nov., G. elisabethjaechae sp.nov, G. valida DELEVE, 1970, G. vietnamensis sp.nov. Habitus and detail drawings of characteristic structures of genitalia and pregenital segments of males are given.

}, keywords = {Coleoptera, Elmidae, Graphelmis, Southeast Asia, taxonomy}, author = {F. {\v C}iampor Jr} } @article {19, title = {Systematic revision of the genus Graphelmis (Coleoptera: Elmidae) V. Graphelmis picta species group}, journal = {Entomological Problems}, volume = {34}, year = {2004}, pages = {55-102}, chapter = {55}, abstract = {

{\v C}IAMPOR Jr., F. \& KODADA, J. 2004. Systematic revision of the genus Graphelmis (Coleoptera: Elmidae) V. Graphelmis picta species group. Entomol. Probl. 34(1\–2): 55\–102. \– The fifth and largest new species group Graphelmis picta is designated within the genus Graphelmis DEL\ÈVE, 1968, comprising nine known and twenty five new species: G. picta (REITTER, 1886), G. biroi (BOLLOW, 1942), G. verpa J\ÄCH, 1985, G. binervosa (REITTER, 1886), G. diaphoroderes J\ÄCH, 1985, G. lomata J\ÄCH, 1985, G. oxytela J\ÄCH, 1985, G. pallidipes (CARTER, 1926), G. palawanensis sp.nov., G. ceylonica (MOTSCHULSKY, 1859), G. loebli sp.nov., G. jendeki sp.nov., G. kubani sp.nov., G. convexa sp.nov., G. balkei sp.nov., G. clava sp.nov., G. riberai sp.nov., G. picea sp.nov., G. vogleri sp.nov., G. basalis sp.nov., G. hlavaci sp.nov., G. securipes sp.nov., G. strbaki sp.nov., G. tuberculata sp.nov., G. philemoni sp.nov., G. togianica sp.nov., G. posoica sp.nov., G. reitteri sp.nov., G. insolita sp.nov., G. darwini sp.nov., G. schneideri sp.nov., G. jesusi sp.nov., G. bruneica sp.nov., G. gibberosa sp.nov. Habitus and detail drawings of characteristic structures of genitalia, pregenital segments and male metatibiae are given.

}, keywords = {Coleoptera, Elmidae, Graphelmis, south-east Asia, taxonomy}, author = {F. {\v C}iampor Jr and J. Kodada} } @article {18, title = {Systematic revision of the genus Graphelmis (Coleoptera: Elmidae) IV. Graphelmis scapularis and Graphelmis clermonti species groups}, journal = {Entomological Problems}, volume = {34}, year = {2004}, pages = {1-20}, chapter = {1}, abstract = {

{\v C}IAMPOR Jr., F. 2004. Systematic revision of the genus Graphelmis (Coleoptera: Elmidae) IV. Graphelmis scapularis and Graphelmis clermonti species groups. Entomol. Probl. 34(1\–2): 1\–20. \– The Graphelmis scapularis and G. clermonti species groups are designated within the genus Graphelmis DEL\ÈVE, 1968. The former includes only one, already known species, the latter comprises three known and six newly described species: G. clermonti (PIC, 1923), G. ambigua DEL\ÈVE, 1970, G. shirahatai (NOMURA, 1958), G. boukali sp.nov., G. spiralis sp.nov., G. schillhammeri sp.nov., G. fulvolineata sp.nov., G. cornuta sp.nov. and G. sausai sp.nov. Diagnoses of the species groups, habitus and detail drawings of characteristic structures of genitalia and pregeni- tal segments and scanning electron microscopy photographs of some male metatibiae are presented.

}, keywords = {Coleoptera, Elmidae, Graphelmis, south-east Asia, taxonomy}, author = {F. {\v C}iampor Jr} } @article {17, title = {Systematic revision of the genus Graphelmis (Coleoptera: Elmidae) III. Graphelmis labralis species group}, journal = {Entomological Problems}, volume = {33}, year = {2003}, pages = {31-44}, chapter = {31}, abstract = {

{\v C}IAMPOR Jr., F. 2003. Systematic revision of the genus Graphelmis (Coleoptera: Elmidae) III. Graphelmis labralis species group. Entomol. Probl. 33(1\–2): 31\– 44. \– The fourth new species group Graphelmis labralis is designated within the genus Graphelmis DEL\ÈVE, 1968, comprising six new species: G. labralis sp.nov., G. tawauica sp.nov., G. kuamutensis sp.nov., G. brezanskae sp.nov., G. kodadai sp.nov., G. temburongensis sp.nov. Habitus and detail drawings of characteristic structures of genitalia and pregenital segments and SEM photographs of male pronotum and metasternal tufts of setae are given.

}, keywords = {Coleoptera, Elmidae, Graphelmis, Southeast Asia, taxonomy}, author = {F. {\v C}iampor Jr} }